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5. 5-833. Marshall. N. B. (1971). ” Cambridge Univ. Press, Cambridge. Martin, J . , Fitzwatcr. S. , and Gordon, R. M. (1900). Iron delicicncy limits phytoplankton growth in Antarctic watcrs. Glohrrl Biogrwchcni. s 4, 5-12. , and Wilson, J. B. (1961). Preliminary field experiments on the relative importance of pressure and tcmpcraturc on the penetration of marine invertebrates into the deep sea. r 12, 302-309. Merrett, N. R . (IW7). A zone of faunal changc in the eastern Atlantic: A response to seasonality in production?

Phytoplankton cells are small and their turnover is rapid, so the standing crop of plant biomass is small and dilute. Moreover, there appear to be only some 5000 species of phytoplankton in the oceans (Tett and Barton, 1995), compared with an estimated 250,000 species of green plants on land. Oceanic herbivores either are suspension feeders or feed on individual particles and so functionally they, too, have to be very small relative to terrestrial herbivores. These grazers are mostly small zooplankton, except in those regions (or seasons) where large diatoms are the dominant primary producers and larger species 1.

Biomass of thc invertebrate megabenthos from 500 to 4100 m in the Northeast Atlantic Ocean. Mrir. Biol. 93, 69-81. , Conkright, M. , Right, J . L.. Najjar. R. , and Mantyla, A. (1993). Distribution of nitrate, phosphate and silicatc in the world oceans. Prog. Ocwnogr. 31, 245-274. Longhurst. A. R. ( I 995). Seasonal cycles of pelagic production and consumption. Prog. Ocermogr. 36, 77-168. Longhurst. A. R.. and Harrison. W. G . (1988). Vcrtical nitrogen flux from the oceanic photic zone by dicl migrant zooplankton and nekton.

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